I don't know how pertinent this info will be, but when I did my volunteer work at wolf park, I learned a lot about wolf behavior. We absolutely had more male wolves in the main pack as opposed to female wolves, because of the rivalry that develops between the females. The lady that you talked with is correct in that "the males will just posture and show teeth, but one will back down while the females will get into a bloody fight". I have experienced this with wolves. Females are very competitive with other females. There was an occasion at the park where we had to remove one of the female wolves because she nearly killed a pack member. Rarely will an alpha female "allow" another female to mate or have pups within a pack. Usually only the alpha pair is the pair that mates and has pups in a pack. Packs are usually extended families. At the park, we did have a very rare occurrance when we had 2 litters of puppies one year. But, after all was said and done, the alpha female took role as mother of all the pups.